I don’t know that’s I use the paper towels. Normally, if they are going to go bad, they mold over within two weeks. This is how I incubate my eggs.
I incubate the eggs the same way that Lynda (kinyonga) does. Put them in a plastic Tupperware shoe box with about 3 inches of moist vermiculite...
They can not lay follicles. This is the surgery performed by my vet when we lived in Charleston, SC.
https://birdsandexotics.com/blog/case-study-3-napoleon-the-chameleon/
How long has it been since the X-ray? She might be egg bound. If the follicles do not harden to eggs, she will not be able to lay them. Sometimes they do absorb the eggs but often they do not.