“Regarding the causes of twinning, scientific articles on wild and captive cases hypothesise aberrant incubation conditions, such as atypical temperatures and humidity, infections, inbreeding or genetic mutations (especially in conjoined twins).”..
https://www.biotaxa.org/hn/article/download/80064/77271
https://www.biotaxa.org/hn/article/download/80064/77271