"The presence of variously sized juveniles with non-gravid and gravid adult females during surveys at the same site suggested that this species might exhibit asynchronous reproduction. We observed a possible mechanism for predator deterrence in this species from repulsive material stored in temporal pouches"...
http://amphibian-reptile-conservati...12_2_[General_Section]_83-89_e160_low_res.pdf
http://amphibian-reptile-conservati...12_2_[General_Section]_83-89_e160_low_res.pdf