A Comparative Perspective on Extra-retinal Photoreception....

[kinyonga]

"Blinded chameleons (species unspecified) have been shown to retain their ability to change body color in response to dermal photostimulation, implying a direct detection of light in the skin"..
https://www.cell.com/trends/endocrinology-metabolism/pdf/S1043-2760(18)30191-7.pdf