“the eggs of non-avian reptiles contain multiple factors to protect the embryo from damage due to microbial infection”…
https://libstore.ugent.be/fulltxt/RUG01/003/008/178/RUG01-003008178_2021_0001_AC.pdf
https://libstore.ugent.be/fulltxt/RUG01/003/008/178/RUG01-003008178_2021_0001_AC.pdf